Chief of Russian intelligence, Sergei Naryshkin, has hinted that Germany was responsible for poisoning Russian opposition leader Alexei Navalny, the BBC reports, without actually making an outright accusation.

Naryshkin asserted that there was "no trace of poison" in Navalny's body when he was transported from Russia for treatment in a German hospital, where he is currently still a patient, and that Russia therefore "has questions" for Germany.

He added that Russia had already destroyed all its reserves of the Novichok nerve agent allegedly used to poison Navalny with.